Following the conclusion of the King of Iron Fist Tournament 4, Heihachi Mishima defeated his son Kazuya Mishima and led him to Hon-Maru Temple, planning to let both Kazuya and his grandson, Jin Kazama, battle and steal their respective "Devil Genes". However, Jin defeats both of his predecessors and flies away, leaving the latter two unconscious. Shortly after, several Jack-4 robots invade the compound, programmed to take Heihachi's life. Initially, Kazuya and Heihachi fight the Jack-4s as a team, but eventually, Kazuya throws Heihachi across the room as soon as he notices that Heihachi is getting tired. Then, Kazuya turns into Devil Kazuya and escapes, leaving Heihachi to deal with the Jack-4s by himself. They quickly overwhelm him and self-destruct, leaving Heihachi seemingly dead.
The explosion, however, revives Heihachi's father Jinpachi Mishima from his 44-year slumber chained up beneath the Temple and proceeds to reclaim the Mishima Zaibatsu. A few months later, the King of Iron Fist Tournament 5 is announced. It turns out that Heihachi, in fact, survived the explosion, and now pledges to find out and "break the neck of the one who started this ridiculous tournament!".
The game follows exactly the same storyline as Tekken 5, only with the additions of two new characters and one returning character; Emilie "Lili" De Rochefort (who seeks to destroy the Mishima Zaibatsu and end her father's financial problem), Sergei Dragunov (a member of Spetsnaz who has been sent to capture Jin, particularly while he is in his Devil form) and Armor King II (after the death of the original Armor King between Tekken 3 and Tekken 4, this new Armor King resurface out of nowhere, with his identity and goals remain a mystery to the player).
Returning Characters
- Jin Kazama (has an alternate "Devil" version)
- Kazuya Mishima
- Heihachi Mishima
- Paul Phoenix
- Marshall Law
- Jack-5
- Nina Williams
- King II
- Yoshimitsu
- Lee Chaolan
- Kuma II
- Wang Jinrei
- Anna Williams
- Ganryu
- Lei Wulong
- Baek Doo San
- Bruce Irvin
- Hwoarang
- Eddy Gordo
- Ling Xiaoyu
- Bryan Fury
- Juilia Chang
- Mokujin
- Panda
- Steve Fox
- Craig Marduk
- Christie Monteiro
- Asuka Kazama
- Feng Wei
- Raven
- Roger Jr.
- Jinpachi Mishima (final boss, unlockable for play in the PS3 version)
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